I am Australian and have flown in and out of Australia many times with firearms. I use Qantas almost exclusively, so cant talk about any other airline. Never had a major issue using Qantas when flying with guns/ ammo. Qantas does require written approval to fly with ammo. But its just a formality - its a letter that the airline issue to you and you need to produce when you check in.
You need to get an import permit from the state police of whichever state you are arriving in, and an export permit from Australian Customs. Its a bit of paper warfare, but not really difficult to get. Unless its changed recently, there are no fees involved with Customs permits, and I believe only Western Australia charges a fee for their state permit.
When you arrive in Australia, checked firearms should be taken by airlines staff directly to customs. You should collect all your checked baggage from the carrousel in the arrival terminal, then go to customs and head to the red coloured "anything to declare" aisle and tell them you have to clear firearms through.. Customs officers will want to see both the import and export permit. They will check the serial no on the forms against the firearm/s, and may look at other items like ammo, but may not look at much else. Its usually a pretty straight forward process. Customs staff have to be specifically trained on this, so sometimes the biggest delay is them finding someone who is allowed to inspect your firearms.
When you depart Australia, you need to go to the customs office
at your point of departure from the country. If at all possible, you want a direct flight out of the country, with no domestic stopovers on the outbound route. Several times I have had to deplane at a domestic terminal, gather all my baggage in the domestic terminal, get on a shuttle bus to the international terminal, then go find the customs office, then recheck all my baggage at the international terminal. The customs officer that clears your paperwork out of the country will then escort you to checkin and stay with you until the firearms are accepted by the airline. Most flights from Aus to USA depart from Sydney. Some from cities like Melbourne or maybe Brisbane may be possible, but will generally only fly a few or less days a week and they change the schedules to/from these cities often.
When booking flights, do not accept anything that stop in asian countries as this will
significantly complicate travelling with firearms. Its not difficult to fly from CA or Tx direct to Aus with Qantas, but I cant speak for other airlines.
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